Brooklyn College, Department of Chemistry, B.S., 1968, Summa cum laude with honors in Chemistry; Yale University, Department of Molecular Biophysics and Biochemistry, M. Phil., 1970, Ph.D., 1972 (Ph.D. Thesis Advisor Prof. Carolyn Slayman); Postdoctoral Fellow, University of Pennsylvania, Johnson Research Foundation, Department of Biophysics and Physical Biochemistry (Prof. Walter Bonner), 1972-1973; Research Associate, Rockefeller University, Laboratory of Cell Biology (Prof. David Luck), 1973-1975; Senior Staff Fellow, National Institute of Mental Health, Laboratory of General and Comparative Biochemistry, 1975-1976; The Edward A. Doisy Department of Biochemistry, Saint Louis University School of Medicine, Assistant Professor, 1976-1979; Associate Professor, 1979-1982; Professor, 1982-1986; Medical Mycology Center, Washington University School Medicine, Barnes and Wohl Hospitals, (Dr. G. Medoff, Chief), 1978 to 1986; Ohio Eminent Scholar in Molecular Genetics, Professor of Molecular Genetics, Biochemistry, and Medical Biochemistry, The Ohio State University, Colleges of Biological Sciences and Medicine, 1986-1997; Director, Institute for Cellular and Molecular Biology, University of Texas at Austin, 1997-2016; Mr. and Mrs. A. Frank Smith Jr. Regents Chair in Molecular Biology, and Professor of Molecular Biosciences, University of Texas at Austin, 1997-present; Professor, Department of Oncology, Dell Medical School, 2018-present.

Service: NIH Molecular Biology Study Section, 1982-1986; Neurospora Policy Committee, 1982-1986; Ohio Science and Engineering Round Table, 1990-1997; ASM Council, 1991-1993; The Academy of Medicine, Engineering, and Science of Texas, Board of Directors, 2010-2012; Scientific Advisory Board, RNA Institute of the University at Albany-State University of New York, 2012-present; Editor, Plasmid, 1987–1998; Editorial Board, Molecular and Cellular Biology (ASM), 1988-2009; Associate Editor, Fungal Genetics and Biology, 1988-2005; Editorial Board, RNA, 1997-present; Editorial Board, Proceedings of the National Academy of Sciences, 2006-2009; Editorial Board, Mobile DNA, 2014-present.

Honors: Member, National Academy of Sciences (2004); Fellow, American Academy of Arts and Sciences (1995); Fellow, American Association for the Advancement of Science (2001); Fellow, American Academy of Microbiology (2004); Member, Academy of Medicine, Engineering, and Science of Texas (2004); Wilbur Cross Medal, Yale University (2013).